Facultad a tiempo completo (Full-Time Faculty)
Ciencias Forenses (Forensic Sciences)
Anya T. Parrilla Díaz, Ph.D.
- Office Address: Natural Sciences Building, University of Puerto Rico at Carolina
- Email: anya.parrilla@upr.edu
- Phone extension: 4555
Courses:
- General Biology I & II, BIOL 3011, 3013
- General Biology Laboratory I & II, BIOL 3013, 3014
- Introduction to Botany Course and Laboratory, BIOL 3417, 3418
- Human Ecology, ESGE 3007
- Fundamentals of Biology I & II, CIBI 3001, 3002
Education:
- Bachelor: S., Biology, University of Puerto Rico, Río Piedras Campus, 1979
- Master: S., Biology, University of Puerto Rico, Río Piedras Campus, 1984
- Doctoral: Philosophy, Applied Ethics, Complutense University of Madrid, Spain, 2013
Honors and Awards:
- Distinguished Scientist (STEM) 2023, University of Puerto Rico, March 2023
Professional Affiliations:
- Advisory Board Member, School of Nursing, University of Puerto Rico Medical Sciences Campus, since 2022
Research:
My greatest research interest is on sustainable development from an ethical and human rights perspective (since 2013). We live a global ecological crisis without precedent and the sustainable development model is an ethical alternative to counteract this crisis. Other research activity: Design and Execute a Recovery Plan for the Community Forest at UPRCA (2018-19). Research proposal approved by the Research and Academic Creation Institutional Committee. Its main purpose was to assess the impact of hurricane María on the flora of the UPRCA Community Forest and to develop a recovery and restoration plan for the preserved area. Development of a Digital Photographic Database of the Puerto Rican Flora. University of Puerto Rico at Carolina, Natural Sciences Department (2001-2005).
Publications:
- Parrilla Díaz, A. T. (2024). El pensamiento ético-ecológico en la visión sostenible del desarrollo. (The ethical-ecological thinking in the sustainable vision of development) Milenio, 23(1), 32–49. Recuperado a partir de https://revistas.upr.edu/index.php/milenio/article/view/21776
- Parrilla, A. T. (2017) El derecho humano a un desarrollo sostenible en solidaridad con la Naturaleza. (The Human Right to Sustainable Development in Solidarity with Nature). Revista Iberoamericana de Bioética, No. 4, 01-11.
- Parrilla, A.T. and J. D. Ackerman. (1990). Epiphyte Roots: anatomical correlates to environmental parameters in Puerto Rican orchids. Orquídea (Mex.) 12 (1):105-116.
Carlos H. Deliz García, Ph.D.
- Office Address: University of Puerto Rico at Carolina Campus
Department of Natural Sciences
2110 Av. Sur, Carolina PR, 00984 - Email: carlos.deliz1@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- General Chemistry I & II, QUIM 3001/QUIM 3002
- Laboratory of General Chemistry I & II, QUIM 3003/QUIM3004
- Organic Chemistry I & II, QUIM 3031/QUIM 3032
- Laboratory Organic Chemistry I & II, QUIM 3033/QUIM 3034
Other Courses:
- Fundamental of General Chemistry, QUIM 3021
- Laboratory of Fundamental of General Chemistry, QUIM 3023
- Fundamental of General Chemistry and Biochemistry, QUIM 3022
- Laboratory of Fundamental of General Chemistry and Biochemistry, QUIM 3024
Education:
- Bachelor: S., Industrial Chemical Process, University of Puerto Rico at Arecibo Campus, 1995.
- Master: S., Applied Chemistry, Pontifical Catholic University of Puerto Rico, 2008.
- Doctoral: D, Environmental Science major on Analytical Physics of Material, University of Turabo, 2018.
Professional Affiliations:
- Colegio de Quimicos de Puerto Rico, 1998
Research:
- Physical chemistry of materials.
Publications:
- Peer-Review Manuscript for International Journal of Environment and Climate Change.
- Efficiency Comparator: Carbon Monoxide Converter Emission by Metallic Osmium Support in Titanium Dioxide and Zirconium Dioxide. Journal of Industrial Pollution Control 37(7) (2021) pp 1-14.
- Carbon Monoxide Converter Emission by Osmium support in Zirconium Dioxide. Journal of Industrial Pollution Control. Journal of Industrial Pollution Control 37(7) (2021) pp 1-13.
- Chapter Book. Chapter 5. Modern Advances in Geography, Environment and Earth Sciences Vol. 1. Evaluating the Effects of Synthesis Methods on the Catalytic Activity of Platinum and Osmium Supported on Titania for Carbon Monoxide Oxidation. Print ISBN: 978-93-90149-92-6, eBook ISBN: 978-93-90149-93-3. oct 2020.
- Alternate Methods for the Calibration of the Dissolution System Apparatus I & II. Publication. Book, 2019. ProQuest. ISBN 9781392810187.
- Effects of the synthesis method on the catalytic activity of platinum and osmium supported on Titania for carbon monoxide oxidation. Journal of Industrial Pollution Control 35(2) (2019) pp. 2283-2293.
- Catalysis in Environmental Application: Carbon Monoxide & Converter Emission. Publication, Library of Congress United States. Book, 2018. TX 8-688-199.
- XV Encuentro de Investigadores Universidad del Turabo, Gurabo PR 2017.
- XIV Encuentro de Investigadores Universidad del Turabo, Gurabo PR 2016.
- 51th Junior Technical Meeting, Pontificia Universidad Católica de PR, Ponce. 2016.
- Convención de Químicos de PR, Plenaria. Tópicos en Químicas. 2008

Edgardo Rivera Tirado, Ph.D.
- Office Address: University of Puerto Rico at Carolina
Department of Natural Sciences
P.O. Box 4800, Carolina, PR 00984-4800 - Office: CN104
- Phone extension: 4551
- Email: edgardo.rivera14@upr.edu
Courses:
- Instrumental Analytical Chemistry, QUIM4015
- Instrumental Analytical Chemistry Laboratory, QUIM4016
- Analytical Chemistry, QUIM3025
- Analytical Chemistry Laboratory, QUIM3026
- General Chemistry I & II, QUIM 3001/QUIM 3002
- Laboratory of General Chemistry I & II, QUIM 3003/QUIM3004
Education:
- Bachelor: S. Industrial Chemistry, Department of Chemistry, University of Puerto Rico at Humacao (1997-2002)
- Doctoral: D., Analytical Chemistry, Department of Chemistry, The University of Akron, OH, USA (2003-2007)
- Post Doctoral: Postdoctoral Research Associate, Measurement Science/Mass Spectrometry, Lubrizol Advanced Materials, Inc., Brecksville, OH, USA (2007)
Honors and Awards:
- Recognition as an American Chemical Society (ACS) Chemistry Ambassador by receiving the Outstanding Chapter Award (2015, 2016), Commendable Chapter Awards (2013, 2014) and the Green Chemistry Student Chapter Awards (2013, 2014, 2015) at the University of Puerto Rico at Cayey.
- Recognition and Dedicatory awards as a Professor of Chemistry and Advisor of the University of Puerto Rico at Cayey Chemistry Student Chapter, respectively (2013 and 2016).
- Poster Second Place Award in the 23rd Annual Lubrizol Corporation Technical Symposium (2007).
- Lubrizol Corporation Scholarship in Chemistry (2005-2007).
- The Dr. Harold G. Cassidy Scholarship in Chemistry, The University of Akron (2006).
- Award for the Launched of “Deracoxib” Product at Pfizer Pharmaceutical Company (2003).
Professional Affiliations:
- Member of the Asociación de Supervisores y Gerenciales de la Universidad de Puerto Rico (A.S.G.U.P.R) (2020-2021).
- Member of the Sociedad de Administradores de Investigación de Puerto Rico (SAI) (2018-2020).
- Member of the American Society for Mass Spectrometry (ASMS) since 2005-current.
- Member of the American Chemistry Society (ACS) since 1997-current.
Research:
- Mass Spectrometry (MS)
- Solid-Phase Microextraction (SPME)
- Headspace Gas Chromatography/Mass Spectrometry (HS-GC/MS)
Publications:
- E. Rivera-Tirado; D. J. Aaserud; C. Wesdemiotis. Characterization of polyisobutylene succinic anhydride chemistries using mass spectrometry. J. Appl. Polym. Sci. 2012, 124, 2682–2690.
- E. Rivera-Tirado; M. López-Casillas; C. Wesdemiotis. Characterization of diazeniumdiolate nitric oxide donors (NONOates) by electrospray ionization mass spectrometry. Rapid Commun. Mass Spectrom. 2011, 25, 3581–3586.
- E. Rivera-Tirado; C. Wesdemiotis. Characterization of polyethylenimine by electrospray ionization and matrix-assisted laser desorption/ionization. J. Mass. Spectrom. 2011, 46, 876–883.
- E. Rivera-Tirado; C. Wesdemiotis. Fragmentation characteristics of Bn (N=2-15) ions from protonated peptides. Rapid Commun. Mass Spectrom. 2011, 25, 2283–2290.

Ernes C Aragonés Geney, Ph.D.
- Office Address: Department of Natural Sciences, University of Puerto Rico
- Email: ernest.aragones@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences:
- Introductory mathematics, MATE 3001
- Precalculus I & II, MATE 3171 / MATE 3172
- Analytic geometry and Precalculus, MATE 3018
- Calculus I, MATE 3031
Other Courses:
– Calculus II, MATE 3032
- Quantitative Methods I & II, MECU 3031 / MECU 3032
Education:
- Bachelor: S., Mathematics, University of Cartagena, 2003
- Master: S., Pure Mathematics, University of Puerto Rico, Mayagüez Campus,2010
- Doctoral: D., Mathematics, University of Puerto Rico, Río Piedras Campus, 2020
Honors and Awards:
– [N/A]
Professional Affiliations:
- American Mathematical Society/Member Since 2025
Research:
My research interests lie in the areas of functional analysis, differential equations, and the controllability of fractional differential systems. I am currently focused on investigating controllability properties and control strategies for systems governed by fractional-order differential equations, with an emphasis on both theoretical analysis and computational approaches. Additionally, I am interested in the broader applications of fractional calculus in modeling complex dynamical systems exhibiting memory and hereditary effects.
Publications:
- E. Aragones, V. Keyantuo and M. Warma, Approximate and Mean Approximate Controllability Properties for Hilfer Time Fractional Differential Equations. Vietnam Journal of Mathematics, 2020)
- E. Aragones, V. Keyantuo and M. Warma, Memory Approximate Controllability Properties for Higher Order Hilfer Time Fractional Evolution Equations. Evolution Equations and Control Theory, 2023

Freisa M. Joaquín Ovalle
- Office Address: Department of Natural Sciences, CN-104, University of Puerto Rico – Carolina
- Email: freisa.joaquinovalle@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- Forensic Toxicology, CIFO4035
- Forensic Toxicology Laboratory, CIFO4036
- Controlled Substances Laboratory, CIFO4046
- General Chemistry I & II, QUIM 3001/QUIM 3002
- Laboratory of General Chemistry I & II, QUIM 3003/QUIM3004
- Organic Chemistry I & II, QUIM 3031/QUIM 3032
- Laboratory Organic Chemistry I & II, QUIM 3033/QUIM 3034
Other Courses:
- Fundamental of General Chemistry, QUIM 3021
- Laboratory of Fundamental of General Chemistry, QUIM 3023
- Fundamental of General Chemistry and Biochemistry, QUIM 3022
- Laboratory of Fundamental of General Chemistry and Biochemistry, QUIM 3024
Education:
- Bachelor: S., Chemistry – University of Puerto Rico Rio Piedras Campus, 2008
- Doctoral: D., Chemistry – University of Puerto Rico, 2022
Professional Affiliations:
- Forensic Science Student Association – Advisor – Since 2023
- American Chemical Society – Regular member
- American Association of Chemistry Teachers – Regular member
- American Society for Biochemistry and Molecular Biology – Early career member
- Biochemical Society – Early career member
Research:
- Focuses on forensic proteomics applications in cancer toxicology
- Interests in protein biomarkers associated with toxicant-induced carcinogenesis
- Identification and quantitation of molecular signatures in biological samples
- Aims to strengthen forensic diagnostics linked to environmental and chemical exposures
- Integrates forensic science with biomedical research for enhanced toxicological interpretation
Publications:
- “Oxidative Stress- and Autophagy-Inducing Effects of PSI-LHCI from Botryococcus braunii in Breast Cancer Cells” Freisa M. Joaquín-Ovalle; Grace Guihurt; Vanessa Barcelo-Bovea; Andraous Hani Saba; Nicole C. Fontanet-Gómez; Josell Ramirez-Paz; Yasuhiro Kashino; Zally Torres-Martinez; Katerina Doble Cacho; Louis J. Delinois; Yamixa Delgado; Kai Griebenow. BioTech 2022, 11(2), 9; https://doi.org/10.3390/biotech11020009
- “Folate-Decorated Cross-Linked Cytochrome c Nanoparticles for Active Targeting of Non-Small Cell Lung Carcinoma (NSCLC)” Irivette Dominguez-Martinez; Freisa M. Joaquín-Ovalle; Yancy Ferrer-Acosta; Kai Griebenow. Pharmaceutics 2022, 14(3), 490; https://doi.org/10.3390/pharmaceutics14030490
- “Key genes and drug delivery systems to improve the efficiency of chemotherapy” Zally Torres- Martinez; Yamixa Delgado; Yancy Ferrer-Acosta; Ivette J. Suarez-Arroyo; Freisa M. Joaquín- Ovalle; Louis J. Delinois; Kai Griebenow. Cancer Drug Resist. (2021); 4: 163-191. http://dx.doi.org/10.20517/cdr.2020.64
- “Optimization and characterization of protein nanoparticles for the targeted and smart delivery of Cytochrome c to non-small cell lung carcinoma” Vanessa Barcelo-Bovea; Irivette Dominguez- Martinez; Freisa M. Joaquín-Ovalle; Luis A. Amador; Elizabeth Castro-Rivera; Kristofer Medina-Alvarez; Anthony McGoron; Kai Griebenow; Yancy Ferrer-Acosta. Cancers (2020), 12(5), 1215. https://doi.org/10.3390/cancers12051215.
- “Thiol-maleimide poly(ethylene glycol) crosslinking of L-asparaginase subunits at recombinant cysteine residues introduced by mutagenesis” Josell Ramirez-Paz; Manoj Saxena; Louis J. Delinois; Freisa M. Joaquín-Ovalle; Shiru Lin; Zhongfang Chen; Virginia A. Rojas-Nieves; Kai Griebenow. PloS One (2018), 13(7): e0197643. https://doi.org/10.1371/journal.pone.0197643.
José A. Cruz-Vega, Ph.D.
- Office Address: Natural Sciences Building, University of Puerto Rico at Carolina
- Phone extension: 4553
- Email: jose.cruz49@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- Introduction to Forensic Sciences, CIFO 3001
- Laboratory of Serology and DNA Technology/ CIFO 4016
- Cellular Biochemistry, BIOL 4545
- General Biology I & II, BIOL 3011, 3013
- General Biology Laboratory I & II, BIOL 3013, 3014
Other Courses:
- Cell & Molecular Biology, BIOL 3907
Education
- Bachelor: S., Biology – University of Puerto Rico at Rio Piedras, 1993
- Doctor: D., Biochemistry – School of Medicine, Medical Sciences Campus, University of Puerto Rico, 1999
Honors and Awards:
N/A
Professional Affiliations:
- American Chemical Society (ACS) member, 2022
Research:
- An unwavering dedication toward updating different scientific issues including modern biotechnology applications, safety and health issues, molecular characterization of diseases, science & technology policies, and Forensic sciences. Excellent ability to transmit scientific information to educate a diverse public.
Publications:
- A. Cruz, R. García, J.F. Rodríguez-Orengo and J.R. Rodríguez-Medina. (2000). Increased Chitin synthesis in response to type II myosin deficiency in Saccharomyce cerevisiae. Mol.Cel.Biol.Res.Comm. 3, 20-25.
- José A. Cruz-Vega, BS; Lilliam Villanueva, MS; José R. Rodríguez-Medina, Ph D. (1998). Flow Cytometry Analysis of Cell Cycle in Myosin II-Deficient Yeast. PRHSJ: 17 (4) 323 326.
- José R. Rodríguez-Medina1, José A. Cruz1, Phillips W. Robbins2, Erfei Bi3, and John R. Pringle3 (1998). Elevated Expression of Chitinase 1 and Chitin Synthesis in Myosin II-deficient Saccharomyces cerevisiae. Cell. Mol. Biol: 44 (6), 919-925.
- Mo-Quen Klinkert *, Francesca Bugli, José Cruz2, Boris Engels, Donato Cioli (1996). Sequence conservation of Schistosome Cyclophilins1. Mol.Biochem. Parasitol:81, 239-242.
José Orlando García Colón, PhD
- Office Address: University of Puerto Rico at Carolina
Natural Sciences Department
PO Box 4800
Carolina, PR 00984 - Phone extension: 4556
- Email: jose.garcia14@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- Introduction to Forensic Sciences, CIFO 3001
- Forensic Toxicology, CIFO 4035
- Forensic Serology and DNA’s Technology Laboratory, CIFO 4016
- Forensic Sciences Capstone Seminar, CIFO 4990
- General Biology I & II, BIOL 3011, 3013
- General Biology Laboratory I & II, BIOL 3013, 3014
Other Courses:
- Human Biology III & IV, BIOL 3791, 3793
- Human Biology Laboratories III & IV, BIOL 3792, 3794
- Animal Organismal Biology, BIOL 3425
- Biology Undergraduate Research, BIOL 3108
- Human Ecology Problems, ESGE 3007
- Fundamentals of Biology I & II, CIBI 3001, 3002
Education:
Bachelor:
- S., General Natural Sciences – University of Puerto Rico, Río Piedras Campus, 2000
Master Courses:
- 32 credits in Pharmaceutical Sciences – University of Puerto Rico, Medical Sciences Campus, 2002
Doctoral:
- D., Medical Physiology – University of Puerto Rico, Medical Sciences Campus, 2010
Post-Doctoral:
- Neurobiology – University of Puerto Rico, Institute of Neurobiology, 2013
Honors and Awards:
– Outstanding Scientific of San Juan Award – Department of Education of Puerto Rico, 2023
– Minority Travel Fellowship Award – American Physiological Society/NIDDK, 2011
– Academic Excellence Award – UPR, Medical Sciences Campus, 2010
– FASEB/MARC Travel Fellowship Award – Federation of American Societies for Experimental Biology, 2009
– Professional Skills Training Workshop Award – American Physiological Society, 2007
– Conference Young Investigator Fellowship Award – International Society for Neurochemistry at Antigua, 2006
– Minority Travel Fellowship Award – American Physiological Society/NIDDK, 2006
– Professional Skills Training Workshop Award – American Physiological Society, 2006
– Research & Travel Award – Center for Pharmaceutical Process Research PR Center, School of Pharmacy, UPR, Medical Sciences Campus, 2001-2002
Professional Affiliations:
– Society for Neurosciences – Regular Member since 2016
– Biophysical Society – Regular Member since 2016
– Biology Honor Society βββ ~ Zeta Mu Chapter – Councilor Member since 2015
– American Physiological Society ~ Puerto Rico Chapter – Founder Directive Board Member since 2009
– American Physiological Society – Regular Member since 2005
– Center for Molecular and Behavioral Neurosciences – CMBN Member since 2004
Research:
Our research objective is to elucidate the role of the interactions between BK potassium channels and membrane rafts in cellular plasma membranes as a critical component for the development of alcohol tolerance. Using a combination of cellular and molecular techniques, we will evaluate the physiological role of the BK channel and its localization in membrane rafts. The subcellular localization of BK channels may provide important insights in our quest to understand the molecular basis of tolerance and its relevance to neuronal plasticity and alcoholism.
Publications:
- Lloret Torres ME, Rivas Jimenez A, Tosado Rodriguez EL, Cardona Jordán K, Lay Rivera XX, Peña Señeriz YL, Capella Muñiz JL, Dieppa Rodríguez M, Ruíz Bolívar DF, González Del Toro, J, Florian Alsina JA, Colón Rivera PA, García Colón JO, Roche-Lima A, Velázque-Marrero C. Comparative mRNA profile analysis from NAc of adolescent male mice after binge-like alcohol exposure eliciting deficits in context fear extinction learning. 2025 Jun; PLoS One. 20(6): e0322576. https://doi.org/10.1371/journal.pone.0322576
- Velázquez-Marrero C, Burgos A, García JO, Palacio S, Marrero HG, Bernardo A, Perez-Lespuir J, Rivera M, Seale GE, and Treistman SN. Molecular alcohol tolerance regulates BK surface expression via Wnt/β-catenin signaling. Journal of Neuroscience. 2016 Oct 12; 36(41): 10625-10639. https://doi.org/10.1523/jneurosci.0491-16.2016
- Salgado IK, Serrano M, García JO, Martinez NA, Maldonado HM, and Silva WI. SorLA in glia: shared subcellular distribution patterns with caveolin-1. Cellular and Molecular Neurobiology. 2011 Nov: 30. doi: https://doi.org/10.1007/s10571-011-9771-5
- García JO. “Capítulo 4: Los Sistemas Inmunitarios, Linfático y Endocrino”. Sciences 6 ~ Textbook. 2009: Santillana Group Inc.; Yabisí Project, 60-77.
- García JO. “Capítulo 2: El Reino Vegetal”. Sciences 5 ~ Textbook. 2009: Santillana Group Inc.; Yabisí Project, 26-43.
- Silva WI, Maldonado HM, Velázquez G, García JO, and González FA. Caveolins in glial cell model systems: from detection to significance. Journal of Neurochemistry. 2007 Nov: 103; 101-12. https://doi.org/10.1111/j.1471-4159.2007.04712.x
Name: José M. Santiago Santana, Ph.D.
Office Address:
- Main office: Natural Sciences Building, 1st Floor, office CN – 005, University of Puerto Rico in Carolina
Neuroscience Research Laboratory:
Natural Sciences Building, 2nd Floor, Laboratory CN -203, University of Puerto Rico in Carolina
Email: jose.santiago13@upr.edu
Phone extension: 4556, 3118
Courses:
Courses on the bachelor’s degree in forensic sciences
– Laboratory of Firearms and Ballistics, CIFO 4056
– Analysis of Controlled Substances, CIFO 4045
- General Biology I & II, BIOL 3011, 3013
- General Biology Laboratory I & II, BIOL 3013, 3014
Other Courses:
- Fundamentals of Biology I & II, CIBI 3001, 3002
- Human Biology III & IV, BIOL 3791, 3793
- Human Biology Laboratories III & IV, BIOL 3792, 3794
- Zoology, BIOL 3425
- Laboratory of Zoology, BIOL 3426
Education:
Bachelor:
- S., Biology with a Major in Human Biology, 2003
University of Puerto Rico, Bayamón Campus
Doctoral:
- D., Physiology and Biophysics, major in Neurosciences, 2011
University of Puerto Rico, Medical Sciences Campus
Honors and Awards:
- National Honor Biology Society, University of Puerto Rico in Carolina, 2016.
- Prize for unconditional support and dedication towards the Student Association.
- National Honor Biology Society, University of Puerto Rico in Carolina, 2015.
- 3rd prize for best counselor of student associations
- Outstanding Research Award, University of Puerto Rico Medical Sciences Campus, 2011.
Professional Affiliations:
– American Academy of Forensic Sciences – Member 2025
– Biophysics Society – Member, 2016-2017
– Society for Neurosciences – Member, 2014-2015
Research:
My work primarily focuses on neuroprotection, neuroregeneration, and the molecular and physiological responses following spinal cord injury (SCI).
Research interests and focus areas:
- Spinal Cord Injury (SCI): Study of cellular and molecular mechanisms involved in SCI, including secondary damage, gliosis, and tissue preservation.
- Hormonal and Pharmacological Therapies: Evaluation of therapeutic potential of tamoxifen and estradiol for neuroprotection and functional recovery post-SCI.
- Oxidative Stress and Neuroinflammation: Studies the impact of these factors in models of SCI and in co-morbid conditions like obesity and stress.
- Behavioral Neuroscience in Animal Models: Analyzes behavioral outcomes, such as fear and anxiety, in rats exposed to obesogenic diets.
- Neurodevelopmental Impact of Diet and Stress: Focused on how adolescent dietary habits and psychogenic stress alter neural and behavioral development.
My laboratory focuses on a multidisciplinary approach by integrating molecular biology, pharmacology, neuroanatomy, and behavioral science to develop potential treatments for traumatic neural conditions.
Publications:
- García-Sánchez, L. M., Colón-Mercado, J. M., Torrado-Tapias, A. I., Padín-Díaz, K., Miranda, J. D., & Santiago-Santana, J. M. (2023). The limited effects of estradiol administration immediately after spinal cord injury. Puerto Rico Health Sciences Journal, 42(1), 23–28.
- Sevillano, M., Vosloo, S., Cotto, I., Dai, Z., Jiang, T., Santiago-Santana, J. M., Padilla, I., Rosario-Pabon, Z., Velez Vega, C., Cordero, J., Alshawabkeh, A., Gu, H., & Pinto, A. (2021). Spatial-temporal targeted and non-targeted surveys to assess microbiological composition of drinking water in Puerto Rico following Hurricane Maria. Water Research X, 13, 100106.
- Santiago-Santana, J. M., Vega-Torres, J. D., Ontiveros-Angel, P., Lee, J. B., Arroyo-Torres, Y., Cruz-González, A., Aponte-Boria, E., Zabala-Ortiz, D., Alvarez Carmona, C., & Figueroa, J. D. (2021). Oxidative stress and neuroinflammation in a rat model of co-morbid obesity and psychogenic stress. Behavioural Brain Research, 400, 113003.
- Vega-Torres, J. D., Haddad, E., Lee, J. B., Kalyan-Masih, P., George, W. I. M., Pérez, L. L., et al. (2018). Exposure to an obesogenic diet during adolescence leads to abnormal maturation of neural and behavioral substrates underpinning fear and anxiety. Brain, Behavior, and Immunity, 70, 96–117.
- Colón, J. M., González, P. A., Cajigas, Á., Maldonado, W. I., Torrado, A. I., Santiago, J. M., et al. (2018). Continuous tamoxifen delivery improves locomotor recovery 6 h after spinal cord injury by neuronal and glial mechanisms in male rats. Experimental Neurology, 299, 109–121.
- Colón, J., Torrado, A., Cajigas, Á., Santiago, J. M., Salgado, I., Arroyo, Y., & Miranda, J. D. (2016). Tamoxifen administration immediately or 24 hours after spinal cord injury improves locomotor recovery and reduces secondary damage in female rats. Journal of Neurotrauma, 33.
- Salgado, I. K., Torrado, A. I., Santiago, J. M., & Miranda, J. D. (2015). Tamoxifen and Src kinase inhibitors as neuroprotective/neuroregenerative drugs after spinal cord injury. Neural Regeneration Research.
- Rosas, O. R., Torrado, A. I., Santiago, J. M., Rodríguez, A., Salgado, I. K., & Miranda, J. D. (2014). Long-term treatment with PP2 after spinal cord injury resulted in functional locomotor recovery and increased spared tissue. Neural Regeneration Research, 24, 2164–2173.
- Mosquera, L., Colón, J. M., Santiago, J. M., Torrado, A. I., Meléndez, M., Segarra, A. C., Rodríguez-Orengo, J. F., & Miranda, J. D. (2014). Tamoxifen and estradiol improved locomotor function and increased spared tissue in rats after spinal cord injury: Their antioxidant effect and role of estrogen receptor alpha. Brain Research, 1561, 11–22.
- Santiago, J. M., Torrado, A. I., Salgado, I. K., König, F., Silva, W. I., & Miranda, J. D. (2013). Expression profile of Flotillin-2 and its possible pathophysiological role after spinal cord injury. Journal of Molecular Neuroscience, 49, 347–359.
- Rodríguez, A. E., Torrado, A. I., Rosas, O. R., Santiago, J. M., Figueroa, J. D., & Miranda, J. D. (2012). Blockade of P2 nucleotide receptors after spinal cord injury reduced the gliotic response and spared tissue. Journal of Molecular Neuroscience, 1, 167–176.
- Arocho, L. C., Figueroa, J. D., Torrado, A. I., Santiago, J. M., Vera, A. E., & Miranda, J. D. (2011). Expression profile and role of ephrinA1 after spinal cord injury. Cellular and Molecular Neurobiology, 7, 1057–1069.
- Rosas, O., Figueroa, J., Torrado, A. I., Rivera, M., Santiago, J. M., König-Toro, F., & Miranda, J. D. (2011). Expression and activation of ephexin is altered in adult rats after spinal cord injury. Developmental Neurobiology, 71, 597–607.
- Santiago, J. M., Rosas, O., Torrado, A. I., González, M. M., Kalyan-Masih, P. O., & Miranda, J. D. (2009). Molecular, anatomical, physiological and behavioral studies of rats treated with buprenorphine. Journal of Neurotrauma, 26, 1783–1793.
Name: Karilys González Nieves, Ph.D.
Office Address:
- Main office: Natural Sciences Building, 1st Floor, office CN-, University of Puerto Rico in Carolina
- Research Laboratory: Natural Sciences Building, 3rd Floor, Laboratory CN -302, University of Puerto Rico in Carolina
- Email: karilys.gonzalez@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- Introduction of Forensic Sciences, CIFO 3001
- Forensic Chemistry, CIFO 3002
- Capstone Seminar, CIFO 4990
General Chemistry I & II, QUIM 3001/QUIM 3002
Laboratory of General Chemistry I & II, QUIM 3003/QUIM3004
Organic Chemistry I & II, QUIM 3031/QUIM 3032
Laboratory Organic Chemistry I & II, QUIM 3033/QUIM 3034
Other Courses:
Fundamental of General Chemistry, QUIM 3021
Laboratory of Fundamental of General Chemistry, QUIM 3023
Fundamental of General Chemistry and Biochemistry, QUIM 3022
Laboratory of Fundamental of General Chemistry and Biochemistry, QUIM 3024
Education:
Bachelor:
S., Chemistry– University of Puerto Rico, Rio Piedras Campus, 2003
Doctoral:
D., Chemistry, major in Inorganic Chemistry – University of Puerto Rico, Rio Piedras Campus, 2013
Honors and Awards:
Osvaldo Ramírez Award, Colegio de Químicos de Puerto Rico, 2025.
Recognition of contributions in science, research, academia, community, and within the Colegio de Químicos de Puerto Rico
Forensic Science Student Society, University of Puerto Rico in Carolina, 2025.
Recognition of academic, mentoring and research contributions
Distinguished Woman Scientist (STEM) 2024, University of Puerto Rico, March 2024
Professional Affiliations:
Puerto Rico Chemistry Association, licensed chemistry (2013- present)
American Chemical Society (ACS) (2003-2022, 2024 at present)
American Association for the Advancement of Science (2018 -201, 2024 at present)
American Association of Chemistry Teachers (AACT) (2015)
Research:
Research interests and focus areas: My research interest is in inorganic chemistry.
Synthesis of metal complexes with electron-rich ligands, with the goal of characterizing them and studying their applications in materials science.
Metal complexes to synthesized Molecular Organic Framework are being explored for applications in batteries.
A new area we are exploring is forensic chemistry, focusing on the study of possible gunshot residue detection.
Publications:
Wentao Hou, Andres Eduardo Araujo-Correa, Shen Qiu, Crystal Otero Velez, Yamna D. Acosta-Tejada, Lexis N. Feliz-Hernández , Karilys González-Nieves, Gerardo Morell, Dalice M. Piñero Cruz, and Xianyong Wu, Electrochemical Studies of Metal Phthalocyanines as Alternative Cathodes for Aqueous Zinc Batteries in “Water-in-Salt” Electrolytes, Batteries 2025, 11, 88.
Vianca Nogué-Guzmán, Alejandro Burgos-Suazo, Javier O. Rivera-Reyes, Vasti P. Montes Quiñones, Paola C. Ramis-Aybar, Adriana C. Burgos-Jiménez, Karilys González-Nieves and Dalice M. Piñero-Cruz, New copper carboxylate pyrene dimers: synthesis, crystal structure, Hirshfeld surface analysis and electrochemical characterization, Acta Crystallographica Section E: Structure Reports Online, 2024, Volume E80, pages 1-9.
Karilys González Nieves, and Dalice Piñero Cruz, Crystal structure of a nickel compound comprising two nickel (II) complexes with different ligand environments: [Ni(tren)(H2O)2][Ni(H2O)6](SO4)2, Acta Crystallographica Section E: Structure Reports Online, 2020, Volume 76, Part 3, pages 314-317.
Konstantinos A. Lazarou, Karilys González Nieves, Indranil Chakrabolty, Raphael G. Raptis, Spontaneous Resolution by Crystallization of an Octanuclear Iron (III) Complex Using Only Racemic Reagents, Chem. Int. Ed. 2019, 58, 7324.
Karilys González, Karinel Nieves and Abimael Rodríguez, 3-chloro-methoxy-N-methylcholest-5-ene-24- carboxamide, Acta Crystallographica, 2012, E68, o3471.
Carlos Jímenez, Karilys González, Abimael Rodríguez, Dolabriferols B and C, non-contiguous polypropionate esters from the tropical sea hare Dolabrifera dolabrifera, Tetrahedron Letters , 2012, 53, 6641–6645.
Karilys González Nieves, 4-((1-(4-bromophenyl)ethylamino)methyl)phenol, Acta Crystallographica, 2011, E67,
Caroline Pérez-Rodríguez, Nashbly Montano, Karilys González and Kai Griebenow, Stabilization of - chymotrypsin at the CH2Cl2/water interface and upon water-in-oil-in-water encapsulation in PLGA microspheres, J. Controlled Release, 2003, 89, 71-85
Name: Maribel Rivera Rivera, MS. EdD.
- Office Address: University of Puerto Rico at Carolina
PO Box 4800
Carolina, PR 00984 - Phone: 787-257-0000 x 4552
- Email: maribel.rivera2@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences:
Introductory mathematics, MATE 3001
Precalculus I & II, MATE 3171 / MATE 3172
Analytic geometry and Precalculus, MATE 3018
Calculus I, MATE 3031
Other Courses:
Calculus II, MATE 3032
Quantitative methods I, MECU 3031
Quantitative methods II, MECU 3032
Introduction to Mathematics I, MATE 3041
Technical mathematics I, MATE 1001
Education:
Bachelor:
S., Secondary education in mathematics, 1993
University of Puerto Rico, Río Piedras Campus
Master:
S., Pure mathematics, 1996
University of Puerto Rico, Río Piedras Campus
Doctoral:
Mathematical education, 2019
University of Puerto Rico, Río Piedras Campus
Name: Maristella Resto Rodríguez, Ph.D.
- Office Address: University of Puerto Rico at Carolina
Natural Sciences Department - Phone extension: 4555
- Email: maristella.resto1@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- General Biology I & II, BIOL 3011/BIOL 3012
- General Biology Laboratory I & II, BIOL 3013/ BIOL 3014
Other Courses:
- General Microbiology, BIOL 3705
Microbiology Laboratory, BIOL 3707
Fundamentals of Biology I & II, CIBI 3001, 3002
Education:
Bachelor:
S., Biology – University of Puerto Rico, Rio Piedras Campus, 1981
Master:
S., Biology with a major in Microbiology) – University of Puerto Rico, Rio Piedras Campus, 1989
Doctoral:
D., Biology, major in Microbiology – University of Puerto Rico Rio Piedras Campus and Medical Sciences Campus, 2000
Associate:
Interior Design- University of Puerto Rico, Carolina Campus, 2010
Honors and Awards:
- Honor certificate for outstanding work University of Puerto Rico Rio Piedras; Dean of Students, 1988
- Distinguished student certificate for having excelled in academic activities as well as in other facets of their university work, University of Puerto Rico Rio Piedras, Natural Sciences Deanery, 1988
- Fogerty Scholarship otorgued for studies in Molecular Parasitology, 1994 Rome, Italy.
Professional Affiliations:
- Society of Microbiologist of Puerto Rico
Research:
Clinical Microbiology, Diagnosis and Molecular techniques, Parasitology
Publications:
Sequence, characterization and localization of Cystein proteinase Cathepsin L in Schistosoma mansoni. Maristella Resto, Anja Michel, Hossam Doneim, Mo-Quen Klinkert, and Wermer Kunz. Molecular and Biochemical Parasitology 73 (1-2): 7-8; Aug 1995.
Name: Naida L. Viera Ruiz, MS
Biology Instructor
- Office Address:
University of Puerto Rico
Carolina Campus
Natural Sciences Department
Biology Area - Phone extension: 4552
- Email: naida.viera@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- General Biology I & II, BIOL 3011, 3013
- General Biology Laboratory I & II, BIOL 3013, 3014
- Genetics, BIOL 3349
Other Courses:
- Biological Sciences I & II, CIBI 3001, CIBI 3002
- General Genetics with Laboratory, BIOL 3305-BIOL 3306
- Fundaments of Microbiology Laboratory, BIOL 3704
Education:
Bachelor:
- S., Biology, University of Puerto Rico, Rio Piedras, 1987
Master:
- S., Biology, University of Puerto Rico, Rio Piedras. 1996
Research interests and focus areas:
Population Genetics
– Genetic variation and structure in natural populations
– Computational approaches for population genetic analysis
Evolutionary Genetics
– Molecular evolution and phylogenetics
– Evolutionary forces shaping genetic diversity
Drosophila Genetics
– Genetic basis of development and behavior in model organisms
– Experimental approaches to study genetic inheritance
Orchids-Fungi Biology
– Mycorrhizal interactions between orchids and fungi
– Symbiotic relationships influencing orchid seed germination and growth
– Fungal biodiversity
Publications:
Bayman, P., Mosquera-Espinosa, A. T., Saladini-Aponte, C. M., Hurtado-Guevara, N. C., & Viera-Ruiz, N. L. (2016). Age-Dependent Mycorrhizal specificity in an invasive orchid, Oeceoclades Maculata. American Journal of Botany, 103(11), 1880-1889. https://doi.org/10.3732/ajb.1600127
Name: Oscar Tomaiconza Ataulluco, Ph.D.
- Office Address:
Department of Natural Sciences
CN-007
University of Puerto Rico at Carolina Campus - Email: oscar.tomaiconza@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
Introductory mathematics, MATE 3001
Precalculus I & II, MATE 3171 / MATE 3172
Analytic geometry and Precalculus, MATE 3018
Calculus I, MATE 3031
Applied Statistics I, ESMA 3101
Other Courses:
Quantitative Methods I & II, MECU 3031 / MECU 3032
Technical Mathematics I, MATE 1001
Introduction to Mathematics I, MATE 3041
Education:
Bachelor:
S., Mathematics, National University of San Antonio Abad of Cusco, Perú, 1999 – 2004
Master:
S., Pure Mathematics, University of Puerto Rico, Mayagüez, 2007 – 2010
A., Economics, University of Puerto Rico, Río Piedras, 2021 – 2025
Doctoral:
D., Mathematics, University of Puerto Rico, Río Piedras, 2013 – 2018
Honors and Awards:
Golden Key International Honor Society, Chapter of the University of Puerto Rico, Río Piedras Campus. 2022
One of the best five positions in academic yield in San Antonio Abad del Cusco of University, 2004
One of the best three positions in academic yield in San Antonio Abad del Cusco of University, 2003
Research:
Research Interests:
Discrete Mathematics
Combinatorics
Algebraic graph theory
Mathematical Economics
Quantitative Methods
Current Projects:
“Manipulación fiscal en años electorales: un análisis del ciclo politico-económico en Puerto Rico (1980-2024)”
The main objective of this research project is to analyze the relationship between fiscal policy and elections, a subject of debate in emerging economies where political cycles can influence state budget decisions. This study focuses on the existence or absence of fiscal manipulation practices during election years in Puerto Rico between 1980 and 2024.
Focus Areas:
Discrete Mathematics and Graph Theory
Dominating Sets in Cayley graphs: Investigated efficient dominating sets in Cayley graphs, focusing on their applications in network design and optimization.
Quantitative Methods and Data Analysis
Optimization and Game Theory: Applied mathematical principles to model and analyze economic behaviors, leveraging optimization techniques advanced statistical techniques.
Publications:
Dejter, I. J., & Tomaiconza, O. (2017). Nonexistence of efficient dominating sets in the Cayley graphs generated by transposition trees of diameter 3. Discrete Applied Mathematics, 232,116-124. https://doi.org/10.1016/j.dam.2017.07.013
Vogel, J. H., Santori-Aymat, M. E., Tomaiconza, O., Cortés-Lumbi., Fernández-Maldonado, M. (2023). Bounded openness over natural information. En J. Martínez-Alier & R. Muradian (Eds.), Elgar Encyclopedia of Ecological Economics (Cap. 6). Edward Elgar Publishing. https://www.elgaronline.com/display/book/9781802200416/ch06.xml
Vogel, J. H., Santori-Aymat, M. E., Tomaiconza, O., Cortés-Lumbi., Fernández-Maldonado, M. (2023). Apertura delimitada sobre información natural. En J. Martínez-Alier & R. Muradian (Eds.), Enciclopedia Elgar de Economía Ecológica (cap. 6, pp. 32-38). Edward Elgar Publishing. https://www.actualidadambiental.pe/wp-content/uploads/2023/10/Entrada-de-Apertura-Delimitada-sobre-Informacion-Nacional-ENCICLOPEDIA-ELGAR.pdf
Name: Rafael Méndez Tejeda, Ph.D.
- Office Address:
Universidad de Puerto Rico en Carolina
Department of Natural Science
P.O. Box. 4800
Carolina. PR. 00984 - Phone extension: 4715
- Email: rafael.mendez@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences:
- General Physics I & II, FISI 3011 /FISI 3012
- Lab. Physics I & II, FISI 3013 / FISI 3014
Education:
Bachelor:
S., Physics, Autonomous University of Santo Domingo, Dominican Republic, 1983
Meteorologist (Class II of the World Meteorological Organization) / National Meteorological Office (ONAMET), Dominican Republic, 1983
Meteorologist (Class II of the World Meteorological Organization) / Meteorological Service of Mexico. Meteorological Service of Mexico, 1984
Master:
- M.S., Physics, Specializing in Radio Astronomy, University of Puerto Rico, Río Piedras Campus, Rio Piedras, Puerto Rico, 1990
Doctoral:
D., Physics, Specializing in Climatology, Complutense University of Madrid, Madrid, Spain.
Post-Doctoral:
Postdoc, University of Florida, Gainesville, Florida. The effect of the El Niño phenomenon in the Caribbean, 2003-2005.
Honors and Awards:
- Senior Researcher in the National Program for Researchers in Science, Technology, and Innovation, Ministry of Education, Science, and Technology of the Dominican Republic.
- Dominican Excellence Award in Puerto Rico in 2018.
Professional Affiliations:
Member Puerto Rico Climate Change (PRCC)
American Meteorological Society- Vice-President Puerto Rico Chapter.
American Geophysical Union (EGU)
Real Sociedad Española de Física
Miembro de la Sociedad Dominicana de Físicos ((SODOFI)
Miembro de la Sociedad Iberoamericana de Física y Química.
Research:
My area of interest is research on climate, climate variability, climate change and meteorological phenomena that impact the Caribbean (tropical cyclones, heat waves, coastal erosion, among others).
Publications:
(5 most recent articles of 75+)
https://scholar.google.com/citations?user=NYXytc0AAAAJ&hl=es
https://www.researchgate.net/profile/Rafael-Mendez-Tejeda
Hernández Ayala, J.J.; Méndez Tejeda, R.; Silvagnoli Santos, F.L.; Villafañe Rolón, N.A.; Martis Cruz, N. Trends in Annual, Seasonal, and Daily Temperature and Its Relation to Climate Change in Puerto Rico. Atmosphere 2025, 16, 737. https://doi.org/10.3390/atmos16060737.
Hernández Ayala, J.J.; Méndez-Tejeda, R. Analyzing Trend in Saharan Dust Concentration and Its Relation to Sargassum Blooms in the Eastern Caribbean. Oceans 2024, 5, 637–646. https://doi.org/10.3390/oceans5030036.
Herrera DA, Cook BI, Fasullo J, Anchukaitis KJ, Alessi M, Martinez CJ, Mendez-Tejeda, Rafael; et al. (2023) Observed changes in hydroclimate attributed to human forcing. PLOS Clim 2(11): e0000303. https://doi.org/10.1371/journal.pclm.0000303
Hernández Ayala, J.J.; Méndez Tejeda, R. Examining the Spatiotemporal Changes in the Annual, Seasonal, and Daily Rainfall Climatology of Puerto Rico. Climate2023, 11, 225. https://doi.org/10.3390/cli11110225.
Santos-Corrada, M.; Méndez Tejeda, R.; Flecha, J. and López, E «An analysis of sustainable consumption practices through the role of consumer behavior in the circular economy». J Consumer Behavior. 2023;1–14.
Name: Yaría Arroyo Torres, Ph.D.
- Office Address:
Main office: Natural Sciences Building, 1st Floor, office CN – 005, University of Puerto Rico in CarolinaResearch Laboratory:
Natural Sciences Building, 2nd Floor, Laboratory CN -203, University of Puerto Rico in Carolina - Email: yaria.arroyo@upr.edu
Courses:
Courses on the bachelor’s degree in forensic sciences
- Firearms and Ballistics Laboratory, CIFO 4056
- General Biology I & II, BIOL 3011, 3013
- General Biology Laboratory I & II, BIOL 3013, 3014
Other Courses:
- Human Biology III & IV, BIOL 3791, 3793
- Human Biology Laboratories III & IV, BIOL 3792, 3794
- Zoology, BIOL 3425
- Laboratory of Zoology, BIOL 3426
- Fundamentals of Biology I & II, CIBI 3001, 3002
Education:
Bachelor:
- S., Biology – University of Puerto Rico-Rio Piedras Campus, May 2006
Doctoral:
- D., Physiology and Biophysics; major in DNA repair and mutations – University of Puerto Rico-Medical Sciences Campus, December 2014
Honors and Awards:
- Second Place: Poster Presentation, 34rd Annual Forum of Research and Education. April 2014
- First Place: Poster Presentation, 4th Annual Puerto Rico Physiological Society. February 2014
Professional Affiliations:
- American Academy of Forensic Sciences- Member 2025
- Forensic Science Student Association (FSSA) at UPR-Carolina Campus-Mentor (January 2024-present)
Research:
My research is centered on understanding the biological mechanisms underlying DNA repair mechanism and neuroprotection as well as the physiological and molecular responses linked to approach/avoidance conflict, fear, and anxiety. Adopting an interdisciplinary perspective, my work integrates molecular biology, neuroanatomy, and behavioral neuroscience to examine neurological trauma and to develop novel therapeutic interventions.
- DNA Repair: Investigating how cells repair DNA damage resulting from traumatic events such approach/avoidance conflict, fear, and anxiety.
- Oxidative Stress and Inflammation: Studying the impact of these processes in approach/avoidance behavioral responses and in comorbid conditions such as obesity and chronic psychosocial stress.
- Behavior in Animal Models: Examining the emotional and behavioral effects—particularly those related to approach/avoidance conflict, fear, and anxiety—in rodent models exposed to high-fat diets and behavioral conflict paradigms.
- Neurodevelopmental Effects of Diet and Stress: Assessing the influence of adolescent dietary habits and psychological stress on neural development and behavioral regulation.
My laboratory is dedicated to advancing DNA repair and translational neuroscience research with the goal of enhancing the understanding, diagnosis, and treatment of traumatic and degenerative disorders of the nervous system.
Publications:
- Sierra, J., Simon, T. B., Abu Hilal, D., Arroyo Torres, Y., Santiago Santana, J. M., & Figueroa, J. D. (2025). Impact of adolescent high-fat diet and psychosocial stress on neuroendocrine stress responses and binge eating behavior in adult male Lewis rats. Hormones and Behavior, 171, 105744. https://doi.org/10.1016/j.yhbeh.2025.105744
- Santiago Santana, J. M., Vega-Torres, J. D., Ontiveros Angel, P., Lee, J. B., Arroyo Torres, Y., Cruz González, A. Y., Aponte Boria, E., Zabala Ortiz, D., Álvarez Carmona, C., & Figueroa, J. D. (2020). Oxidative stress and neuroinflammation in a rat model of co-morbid obesity and psychogenic stress. Behavioural Brain Research, 400, 112995. https://doi.org/10.1016/j.bbr.2020.112995
- Vega-Torres, J. D., Haddad, E., Lee, J. B., Kalyan-Masih, P., Maldonado George, W., López Pérez, L., Piñero Vázquez, D. M., Arroyo Torres, Y., Santiago Santana, J. M., Obenaus, A., & Figueroa, J. D. (2018). Exposure to an obesogenic diet during adolescence leads to abnormal maturation of neural and behavioral substrates underpinning fear and anxiety. Behavioural Brain Research, 70, 96–117. https://doi.org/10.1016/j.bbi.2018.01.011
- Colón, J. M., Torrado, A. I., Cajigas, Á., Santiago Santana, J. M., Salgado, I. K., Arroyo Torres, Y., & Miranda, J. D. (2016). Tamoxifen administration immediately or 24 hours after spinal cord injury improves locomotor recovery and reduces secondary damage in female rats. Journal of Neurotrauma, 33(18), 1696–1708. https://doi.org/10.1089/neu.2015.4111(ncbi.nlm.nih.gov)
- Santiago, J. M., Torrado, A. I., Arocho, L. C., Rosas, O. R., Rodríguez, A. E., König Toro, F., Salgado, I. K., Arroyo Torres, Y., Silva, W. I., & Miranda, J. D. (2012). Expression profile of flotillin-2 and its pathophysiological role after spinal cord injury. Journal of Molecular Neuroscience, 49(2), 347–359. https://doi.org/10.1007/s12031-012-9873-7 (link.springer.com











